It was a fairly quiet morning this morning with the only highlight being another Firecrest. With a bit of time to spare I thought it was worth mentioning a few of the more interesting things we've caught over the last week. First up, the (so far only) Pied Flycatcher.
Pied Flycatchers (of both age classes) undergo a partial pre-breeding moult before they leave their wintering grounds. Critically, this means that
both ages show a moult limit in the greater coverts in spring. Not all of the body feathers are moulted during this pre-breeding moult; the result is blotchy brown patches on the neck, scapulars and back that contrast with the black breeding plumage. The autumn/winter plumage of both young and adult Pied Flycatchers is brown, so this contrast is between brown and black plumage not between faded/bleached and new plumage.
The moult limit on all birds can make ageing difficult at times and one has to assess the wear, colour, etc of the feathers that have been retained from the previous autumn's moult (primaries, secondaries, tail, unmoulted greater coverts). This week's bird was reasonably easy to age: the primaries and secondaries are dark (rather than brownish) and in good condition, and the unmoulted outer three greater coverts are almost as black as the moulted pre-breeding greater coverts, indicating an adult (3+cy).
This bird shows a few interesting quirks. Firstly, the greater coverts.
GC10-6 (innermost) are pre-breeding, as is GC4, while GC1-3 (outermost) are post-breeding (from last autumn). GC5 is conspicuously and you could be forgiven for thinking there are in fact three generations of feathers in the greater coverts. GC5 is, however, probably not as interesting as it first seems. Many adult male Pied Flycatchers show a pseudo-moult limit in the greater coverts with the inner 6 to 8 being brown and the outer 2 to 4 being black. This is likely an adaptation to maximise the extent of black in the breeding plumage or to reduce wear on the the feathers that likely be retained for a full year (i.e. not moulted in the pre-breeding) since blacker feather = more melanin = less wear. Probably both factors come into play. GC5 should therefore, despite the apparent differences, be the same generation as GC1-3 (autumn 2012). The moult pattern was the same on both wings.
Examining the right wing further, it's apparent that the bird has retained S6 (the innermost secondary).
Although clearly older than S1-5, this feather is in good condition and is rather dark and black. According to data in Jenni & Winkler
Moult and Ageing of European Passerines retained secondaries in autumn are not uncommon; the number in spring is a lot less. There might be several factors at play (perhaps the moult is suspended rather than arrested and hence completed on the wintering grounds; perhaps birds with retained secondaries are physically less fit and thus more likely to die on migration) but it's probably because of the extensive pre-breeding moult (often including the inner secondary/ies) that we don't notice it more often in spring — it's been 'covered over'.
It's hard to know exactly where the retained S6 on the right wing came from. The most likely answer is that it's a spring 2012 pre-breeding feather that wasn't moulted during the 2012 complete autumn moult.
Checking the left wing, things become more intriguing still. S6 is retained here as well; but it's clearly older than S6 on the right wing!
The conclusion is that this bird shows
four generations of feathers: 2013 pre-breeding; 2012 post-breeding; 2012 pre-breeding; and S6 on the left wing, which must be from autumn 2011. The question then is, what can we say about S6 on the left wing? Despite the fact it's pretty shagged, it's still in reasonably good condition for a feather that must be 18 months old. It's still quite dark and, although the tip has broken off, the feather wear isn't *that* heavy (though still considerably more so than on S6 on the right wing). I'd expect a juvenile feather to be a lot more worn than this, and a lot more faded — indeed, it probably wouldn't be much more than a feather shaft and a few tufts of sandy-brown barbs. If this feather isn't a juvenile feather than it must be an adult feather from the 2011 post-breeding moult; that means this bird should be a 4+cy.
Amongst the huge numbers of Chaffinch we caught the other week there were two interesting birds; they had also retained some feathers that they shouldn't have done. This female had retained some greater coverts on the right wing only (GC1, 4-5 & 9), all three tertials on the right wing and T3 on the left wing, and S5-6 on the right wing.
The retained feathers are in good condition, not too worn, and, in the case of the tertials and secondaries, broad and square-tipped: this indicated that they are adult or adult-type feathers. The tertials and the greater coverts could have been moulted during the post-juvenile moult and would then not allow us to differentiate a 3cy from a 4+cy. However, a post-juvenile moult would not include the secondaries; thus the retained S5-6 allow us to age this bird as a 4+cy.
The male-like colours appearing on the back and breast also support the theory that this is an older bird.
In the second case, an adult male Chaffinch had moulted everything except for the tertials on both wings. The tertials, in contrast to those on the bird above, a narrow and worn; they appear to be juvenile feathers.
It's slightly off-putting that T1 is blacker than T2-3. Perhaps this is simply due to being more protected. If the tertials are juvenile feathers, that would make this bird a 3cy.
Always interesting, we've been catching plenty of Dunnocks the last couple of weeks — in fact, with 227 ringed so far, 2013 is the second-best spring on record. Perfect for testing out a paper that Björn Malmhagen and I have just had accepted for publication in
Ringing & Migration: Ageing Dunnocks
Prunella modularis using plumage characteristics. It seems to work well... though I guess we would say that, wouldn't we?!
Second-year female. Bootiful.